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Phloem is a conduction tissue in plants. The main fuction

conduct food substances such as sugars, amino acids,

micronutrients, lipids etc. from leaves to root and stem.

Phloem may be classified as

primary

or

secondary

on the

basis of its time of appearance in relation to the development

of the plant or organ as a whole.

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Phloem is composed of four kinds of cells

a) Sieve elements

b) Companion cells

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Sieve elements

Sieve elements are main component of phloem. their walls of areas (sieve areas) penetrated by pores.

Sieve elements are divided into on sieve tubes and sieve cells on the basis of specialization into cells hape:

Sieve tubes

Sieve tubes are long tube like structure and are formed from a row of cells arranged in longitudinal series. The end wall of sieve tube cells shows perforation called as sieve plate which establishes the connectivity with the

neighboring sieve cell. The protoplasts of sieve-tube elements contain P-protein (phloem protein or formerly called as slime). P Proteins (Phloem Proteins) are a category of proteins present in the sap of the sieve tubes in

Angiosperm plants. The unique carbohydrate known as callose is associated with conducting sieve-tube elements and it is deposited there in response to mechanical injury, some kind of stimulation. Callose normally accumulates at sieve plates. Callose apparently plays a role in sieve-pore development.

Sieve cells

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Companion cells

• Companion cells and sieve-tube elements are closely

related ontogenetically and occured from the same

mother cell. The companion cell cytoplasm has a

evident nucleus and is connected to sieve tubes

through pits found in lateral walls in Angiospermae.

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Phloem parenchyma

• They are living and have primary wall and simple

pits. Main fuction of phloem parenchyma store

starch, oil and the other ergastic substances like

xylar parenchyma.

• The parenchyma cell in secondary phloem are

two types, phloem parenchyma and ray cells.

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Phloem fibres

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